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The divergence of these species is similar in time to the radiation of the Leuciscinae supposedly centred in Siberia based on fossil records. (sic) from the Lower Miocene of Saudi Arabia showing an early date for the entry of cyprinids to the Afro-Arabian Plate.

Siberia was probably an important dispersion centre for both Leucicinae and Cyprininae at that time. Some species may enter brackish water but the family is primarily a freshwater one.

Other Middle Eastern cyprinid genera are regarded by Durand et al.

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Collares-Pereira (1994) argues that the polyploid condition (e.g. There are 2-4 unbranched rays (including rudimentary ones) in the dorsal and anal fins followed by the more numerous branched rays (the last two branched rays are counted as one).Previously, I completed my undergraduate study with a first class MMath Mathematics degree from the University of Exeter in 2014.Introduction This family contains by far the most species in the Iranian freshwater ichthyofauna and is divided into two files Abramis to Cyprinus (here), and Garra to Vimba (see both in Contents).This is not in general use at this writing and the Cyprinidae is retained as a single family here. (2002) using cytochrome b DNA of Cyprinidae conclude that the the Middle East is an important interchange area for this freshwater ichthyofauna rather than a centre of speciation.The Middle East leuciscine cyprinids have Europe as an important Palearctic influence consistent with the Lago Mare dispersion while the the cyprinine cyprinids show three highly divergent lineages, namely one shared with the Euro-Mediterranean area (Barbus/Luciobarbus), a relict of the Lago Mare dispersion, one shared with Africa (Carasobarbus/Varicorhinus subgenus) and one with Asia (Garra).

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